01 - Preheat oven to 375°F and line a 12-cup muffin tin with paper liners or grease lightly.
02 - Whisk together flour, poppy seeds, baking powder, baking soda, and salt in a large bowl.
03 - In a separate bowl, rub granulated sugar and lemon zest with fingers to release oils.
04 - Add eggs, Greek yogurt, vegetable oil, lemon juice, and vanilla extract to sugar mixture; whisk until smooth.
05 - Pour wet ingredients into dry ingredients and stir gently until just combined, avoiding overmixing.
06 - Divide batter evenly among muffin cups, filling each about three-quarters full.
07 - Bake for 18 to 20 minutes or until a toothpick inserted into the center comes out clean.
08 - Allow muffins to cool in the pan for 5 minutes before transferring to a wire rack to cool completely.